Heteropogon contortus
Poaceae or Grass family
Synonyms: Andropogon contortus
Heteropogon contortus
Poaceae or Grass family
Synonyms: Andropogon contortus
Pili
Lule
Pili grass, Spear grass, Tanglehead, Twisted beardgrass
Natural Range & Environment
Widely distributed throughout the tropics; in Hawaiʻi, an indigenous grass occurring on dry rocky cliffs, ledges, or slopes close to ocean exposure from sea level to 700 m (2,295 ft) on all the main Hawaiian islands. Pili is not as common today as it was in the past and is disappearing in some areas, especially on Oʻahu and Molokaʻi [Wagner et al. 1990].
Pili flowers. Notice in the photo of barbs with attached seeds (right), as well as seeds with no barbs attached (on right). Clos-eup of leaves and seeds, and planting en masse on a college campus.
Description
Heteropogon contortus are perennial. The culms (stems) can grow from 15 (4 dm) to about 40 inches (10 dm) tall with often with glaucus or pale blue-green leaves. The twisted, sharp barbed spikelets attach to seeds. Pili form patches from 60-150 cm (2-5 ft) wide.
Growth Requirements
General
Pili does well in containers of almost any size, but will quickly outgrow smaller ones. For container plants, small applications of foliar feeding with an organic or soluble commercial fertilizer monthly or every other month, from one-third to one-fourth of the recommended strength, is beneficial.
However, pili does better and will grow to its full extent when planted in the ground, where it does not require much care once established. The soil must be well-drained with sand, cinder or organic matter. Pili will tolerate limited partial sun but does best in full sun with good air movement. It can tolerate wet or dry conditions, but overall looks better when given water regularly. It is drought, wind, heat, and salt spray tolerant.
If pili is planted close together, it forms a tall groundcover or dense low hedge. While it does not need to be trimmed, this can be done once a year or every other year, down to six inches to encourage new growth and maintain a clean appearance by removing the tangled masses of seeds and harpoons. Interestingly, when rain or water drops hit the tangled mass, they start to twist and writhe. Keiki (children) love animated things, even animated plants like Sleeping grass (Mimosa pudica) and Venus Fly Traps. You can carefully gather a bunch of pili seeds/harpoons and show your keiki what happens when water drops on lands on it. The seeds/harpoons will twist and curl, seemingly coming alive, just as the vernacular name Twisted beardgrass suggests! (See Etymology).
It is important to use native grasses such as pili in the landscape because they are being displaced by alien grasses and shrubs in wild places. Additionally, grasses and sedges add another shape or form to the landscape, fufilling our natural desire to see more than just broad-leaved plants.
Pests and Diseases
Pili is occasionally subject to attacks by scale insects, mealybugs, and grasshoppers, but is otherwise generally pest-free. Refer to the Pests & Diseases page for treatments.
Uses
Ancient Hawaiian
Early Hawaiians preferred using pili for thatching roofs due to its brown color, neat appearance, and pleasant odor [Abbott 1992, Neal 1965] . Pili was harvested by uprooting a bunch, trimming away the soil, roots, and flowering spikes, and then tying the pili bunch in rows with stems up. The bunches were placed close together, working from the bottom of the frame upwards. Pili thatch needed to be replaced every four to five years. A black dye was created from charcoaled leaf blades [Krauss 1993]. In addition to dye and construction, pili leaves were utilized for stuffing mattresses, padding floors, and tinder [Lincoln 2009]. Medicinally, burned pili and ashes mixed with coconut were used for treating ʻea (thrush) and pāʻaoʻao (childhood disease causing physical weakening) [Handy 1972].
Modern
Both the upright and prostrate forms of pili are cultivated in Hawaiʻi for private and public landscapes, often for cultural displays such as botanical gardens and college campuses. Pili has played a crucial role in conservation efforts on Kahoʻolawe and is also utilized as a roadside grass to combat invasive species along local transportation corridors.
Special Features and Information
General
Heteropogon contortus, a member of the Grass family (Poaceae), is part of a diverse group of plants that includes approximately 12,000 species and 780 genera worldwide. In the Hawaiian Islands, there are over 80 species of native grasses, many of which are endemic, and some are considered rare and endangered [Flora of the Hawaiian Islands].
Pili is believed to be indigenous to the Hawaiian Islands. The seeds of pili have harpoon-like attachments that allow them to attach to fur and feathers for dispersal.
Botanist Joel Lau has identified two forms of pili in Hawaiʻi—an erect type and a prostrate type. The erect type of H. contortus can be found on all the main Hawaiian Islands, while the prostrate type is limited to areas where the trade winds are particularly strong, such as specific sites in northwestern Molokaʻi, northwestern Lānaʻi, and Kalae (South Point) on Hawai`i Island.
Pili serves as an important habitat for various native creatures, including land snails, and its presence in the Hawaiian Islands contributes to the biodiversity and ecological balance of the region. Small terrestrial snails, including Cookeconcha undescribed sp.?, Lyropupa perlonga, and Endodonta undescribed sp.?, were previously abundant among pili at Koko Head, Oʻahu. They are now only known from sub-fossil/fossil sediments at Barbers Point (Kalaeloa), Oʻahu. It is believed that these snails may have also inhabited habitats similar to the bunchgrass (Eragrostis sp.) found on Nīhoa [Christensen & Kirch 1986].
Hawaiian Name
Pili means "to cling or stick" [Hawaiian Dictionaries].
Etymology
The generic name Heteropogon is derived from the Greek heteros, different, and pogon, beard, in reference to the two kinds of spikelets found in the inflorescence of this species. Literally, "varying-beard," (the twisting awns of the female spikelets) [Gledhill 2008]. True to the Latin specific epithet contortus, pili seeds have an unusual habit of contorting when wet to burrow into the soil [Gledhill 2008].
DWE